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The study of beetles; Some skeletal peculiarities of the adults; Internal structures of the adults; Some structural features of larvae and pupae; Food, digestion and the alimentary canal; Blood, osmoregulation, reserves, excretion and endocrine organs; Locomotion, respiration and energetics; The senses; Cuticular properties, appearance, colour and luminosity; Adult and larval behaviour; Development and life-cycles; Cytology and genetics; Water beetles; Special habitats; predation and defence; Symbiotic and parasitic relations; An ecological triangle: beetles, fungi and trees; Herbivorous beetles; Geographical distribution and conservation: beetles as ecological indicators; Evolutionary history of beetles.
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